User:Alandmanson/South African Scarabaeinae
Traditional tribes:
Ateuchini (previously Dichotomiini) -
Coprini -
Deltochilini (Canthonini) -
Eucraniini -
Gymnopleurini -
Oniticellini (including Eurysternina) -
Onitini -
Onthophagini -
Phanaeini -
Scarabaeini -
Sisyphini
source: Bouchard et al. (2011) [1]
Groups of Tarasov (2016): "Basal Scarabaeinae" - Scarabaeini - Coprini sensu novo - Australasian endemic genera - Ateuchini: Scatimina partim - Dichotomiini sensu novo - Ateuchini partim - Phanaeini - Eucranini - Deltochilini (Canthonini) sensu novo - Gymnopleurini - Eurysternini - Sisyphini sensu novo - Onotini - Onthophagini + Oniticellini
Tribe: Traditional Ateuchini (Dichotomiini)
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/0/0a/Sarophorus_costatus_001065-3.jpg/220px-Sarophorus_costatus_001065-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1065 Category:Sarophorus costatus
Tarasov (2016): Sarophorus included in "Basal Scarabaeinae"
Tribe: Scarabaeini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/b/b1/Pachylomera_femoralis_000215-1.jpg/220px-Pachylomera_femoralis_000215-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-215 Category:Pachylomera femoralis
![](https://upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Scarabaeus_caffer_000360-1.jpg/220px-Scarabaeus_caffer_000360-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-360 Category:Scarabaeus caffer
![](https://upload.wikimedia.org/wikipedia/commons/thumb/2/22/Scarabaeus_geminogalenus_000716-1.jpg/220px-Scarabaeus_geminogalenus_000716-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-716 Category:Scarabaeus geminogalenus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/7/7b/Scarabaeus_nigroaeneus_248-1_2016_01_01.jpg/220px-Scarabaeus_nigroaeneus_248-1_2016_01_01.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-248 Category:Scarabaeus nigroaeneus
Tribe: Traditional Deltochilini (Canthonini)
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/b/bd/Epirinus_validus_000916-2.jpg/220px-Epirinus_validus_000916-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-916 Category:Epirinus validus
Tarasov (2016): Epirinus included in Sisyphini sensu novo
Tribe: Traditional Coprini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/0/0a/Copris_bootes_001139-3.jpg/220px-Copris_bootes_001139-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1139 Category:Copris bootes
![](https://upload.wikimedia.org/wikipedia/commons/thumb/4/4e/Copris_fidius_001249-2.jpg/220px-Copris_fidius_001249-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1249 Category:Copris fidius
![](https://upload.wikimedia.org/wikipedia/commons/thumb/8/84/Copris_denticulatus_001143-3.jpg/220px-Copris_denticulatus_001143-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1143 Category:Copris denticulatus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/b/b3/Heliocopris_hamadryas_001146-3.jpg/220px-Heliocopris_hamadryas_001146-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1146 Category:Heliocopris hamadryas
![](https://upload.wikimedia.org/wikipedia/commons/thumb/d/dd/Heliocopris_neptunus_001144-2.jpg/220px-Heliocopris_neptunus_001144-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1144 Category:Heliocopris neptunus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/e/e9/Metacatharsius_opacus_000724-2.jpg/220px-Metacatharsius_opacus_000724-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-724 Category:Metacatharsius opacus
Tarasov (2016): Metacatharsius + Catharsius are sister to a group including Gymnopleurini, Eurysternini & Sisyphini sensu novo
Tribe: Gymnopleurini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Allogymnopleurus_thalassinus_000869-1.jpg/220px-Allogymnopleurus_thalassinus_000869-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-869 Category:Allogymnopleurus thalassinus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Garreta_unicolor_2016_01_01_11_000244-1.jpg/220px-Garreta_unicolor_2016_01_01_11_000244-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-244 Category:Garreta unicolor
![](https://upload.wikimedia.org/wikipedia/commons/thumb/7/7a/Garreta_nitens_246-1_2016_01_01.jpg/220px-Garreta_nitens_246-1_2016_01_01.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-246 Category:Garreta nitens
![](https://upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Gymnopleurus_asperrimus_001205-3.jpg/220px-Gymnopleurus_asperrimus_001205-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1205 Category:Gymnopleurus asperrimus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Gymnopleurus_virens_781-2.jpg/220px-Gymnopleurus_virens_781-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-781 Category:Gymnopleurus virens
Tribe: Sisyphini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Sisyphus_fasciculatus_275-1.jpg/220px-Sisyphus_fasciculatus_275-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-275 Category:Sisyphus fasciculatus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/5/55/Sisyphus_sordidus_864-2.jpg/220px-Sisyphus_sordidus_864-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-864 Category:Sisyphus sordidus
Tribe: Oniticellini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/4/4d/Euoniticellus_africanus_875.jpg/220px-Euoniticellus_africanus_875.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-875 Category:Euoniticellus africanus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/6/6b/Euoniticellus_intermedius_000467-1.jpg/220px-Euoniticellus_intermedius_000467-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-467 Category:Euoniticellus intermedius
![](https://upload.wikimedia.org/wikipedia/commons/thumb/c/c6/Liatongus_militaris_2016_11_19_6712.jpg/220px-Liatongus_militaris_2016_11_19_6712.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-862 Category:Liatongus militaris
![](https://upload.wikimedia.org/wikipedia/commons/thumb/2/24/Oniticellus_pictus_001052-2.jpg/220px-Oniticellus_pictus_001052-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1052 Category:Oniticellus pictus
Tribe: Onthophagini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Digitonthophagus_gazella_001206-3.jpg/220px-Digitonthophagus_gazella_001206-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1206 Category:Digitonthophagus gazella
![](https://upload.wikimedia.org/wikipedia/commons/thumb/0/0c/Onthophagus_alcyonides_000876-3.jpg/220px-Onthophagus_alcyonides_000876-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-876 Category:Onthophagus alcyonides
![](https://upload.wikimedia.org/wikipedia/commons/thumb/a/a8/Onthophagus_aequepubens_000857-3.jpg/220px-Onthophagus_aequepubens_000857-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-857 Category:Onthophagus aequepubens
![](https://upload.wikimedia.org/wikipedia/commons/thumb/0/06/Onthophagus_aeruginosus_000855-1.jpg/220px-Onthophagus_aeruginosus_000855-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-855 Category:Onthophagus aeruginosus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/7/73/Onthophagus_binodis_001239-3.jpg/220px-Onthophagus_binodis_001239-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1239 Category:Onthophagus binodis
![](https://upload.wikimedia.org/wikipedia/commons/thumb/3/3b/Phalops_flavocinctus_000866-1.jpg/220px-Phalops_flavocinctus_000866-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-866 Category:Phalops flavocinctus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/5/5b/Proagoderus_aciculatus_000730-1.jpg/220px-Proagoderus_aciculatus_000730-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-730 Category:Proagoderus aciculatus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Proagoderus_chalcostolus_000872-3.jpg/220px-Proagoderus_chalcostolus_000872-3.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-872 Category:Proagoderus chalcostolus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Proagoderus_lanista_000861-1.jpg/220px-Proagoderus_lanista_000861-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-861 Category:Proagoderus lanista
Tribe: Onitini
[edit]![](https://upload.wikimedia.org/wikipedia/commons/thumb/7/79/Onitis_perpunctatus_001236-1.jpg/220px-Onitis_perpunctatus_001236-1.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1236 Category:Onitis perpunctatus
![](https://upload.wikimedia.org/wikipedia/commons/thumb/3/32/Onitis_tortuosus_001251-2.jpg/220px-Onitis_tortuosus_001251-2.jpg)
http://vmus.adu.org.za/?vm=DungBeetleMAP-1251 Category:Onitis tortuosus
Links
[edit]Dung Beetles of Manyoni Private Game Reserve
DungBeetleMAP
[edit]33 species found for locus = 2930
Caccobius obtusus Copris jacchoides Copris obesus Cyptochirus ambiguus Epirinus relictus
Litocopris simplex Nebulasilvius insularis
Oniticellus planatus Onitis pecuarius
Onthophagus bicavifrons Onthophagus cretus Onthophagus cribripennis Onthophagus deterrens Onthophagus pilosus Onthophagus vigens Onthophagus vinctus
Parvuhowdenius harrisoni
Xinidium dentilabris Xinidium dewitzi